The species scale serves as a UV cutoff in the gravitational sector of an EFT and can depend on the moduli of the theory. In this talk I will demonstrate how in certain setups we can explicitly compute the species scale at any point in field space and will discuss general bounds on the variation of the species scale as a function of the scalar fields valid everywhere in field space. As an application of this, I will show how the species scale constrains the maximum diameter of the field space and, using the emergent string conjecture, will recover the bound on scalar potentials previously found in the context of the TransPlanckian Censorship Conjecture.